The genes that control PEPOEFL transmitted to the descendants were mainly based on additive effect , but also with the non- additive effect .

控制非旗叶光合效应的基因作用以加性效应为主，同时存在非加性效应；

The result showed that the genes that control protein content transmitted to the descendants were mainly based on additive effect, but also with the non-additive effect.

结果表明，控制蛋白质含量的基因作用以加性效应为主，同时存在非加性效应；